Taxacom: Question to botanical experts RE ICNafp Code - rank changes for descriptive names, with changed termination

Fri Mar 31 19:02:32 CDT 2023

Hi Tony,
       "An alternative model for the earliest evolution of vascular plants" published in 2019 (https://nam10.safelinks.protection.outlook.com/?url=https%3A%2F%2Fdoi.org%2F10.1111%2Flet.12323&data=05%7C01%7Ctaxacom%40lists.ku.edu%7C89064c749dd74fe7bac008db32446561%7C3c176536afe643f5b96636feabbe3c1a%7C0%7C0%7C638159041585165897%7CUnknown%7CTWFpbGZsb3d8eyJWIjoiMC4wLjAwMDAiLCJQIjoiV2luMzIiLCJBTiI6Ik1haWwiLCJXVCI6Mn0%3D%7C3000%7C%7C%7C&sdata=Nq09syxWwner%2Flm0VFxLoe1sJpgJ5up205E6tNxyfIc%3D&reserved=0) says: "Protracheophytes include the Horneophytopsida and Aglaophyton/Teruelia."  And the paper proposing the name Eophytidae shows eophytes splitting off between Horneophytopsida and Aglaophyton.  And the wikipedia article on Polysporangiophytes (https://nam10.safelinks.protection.outlook.com/?url=https%3A%2F%2Fen.wikipedia.org%2Fwiki%2FPolysporangiophyte&data=05%7C01%7Ctaxacom%40lists.ku.edu%7C89064c749dd74fe7bac008db32446561%7C3c176536afe643f5b96636feabbe3c1a%7C0%7C0%7C638159041585165897%7CUnknown%7CTWFpbGZsb3d8eyJWIjoiMC4wLjAwMDAiLCJQIjoiV2luMzIiLCJBTiI6Ik1haWwiLCJXVCI6Mn0%3D%7C3000%7C%7C%7C&sdata=zgfsbjTud6vVEG8AtySpjRCOS9PQKD3%2BVPj46svzNbI%3D&reserved=0) shows an older cladogram also showing Horneophytopsida and Aglaophyton as protracheophytes.
                         -----------------Ken

________________________________
From: Tony Rees <tonyrees49 at gmail.com>
Sent: Friday, March 31, 2023 6:20 PM
To: Kenneth Kinman <kinman at hotmail.com>
Cc: taxacom <taxacom at mailman.nhm.ku.edu>
Subject: Re: Taxacom: Question to botanical experts RE ICNafp Code - rank changes for descriptive names, with changed termination

Hi Ken, that is an interesting suggestion, that I would have to look further into before formulating a response - but others (more expert than me) are very welcome to comment.

FWIW my current IRMNG remarks for Aglaophyton read:
"Originally allocated to fam. Rhyniaceae, but considered distinct from that group by later authors. Subsequently identified as the sporophyte of the earlier described Lyonophyton W. Remy & R. Remy, 1980; the two names can co-exist as separate form-genera, although Novikoff & Barabasz-Krasny, 2015 treat Aglaophyton as a synonym of Lyonophyton, and place the latter in its own family Lyonophytaceae Doweld."

Presently IRMNG has Aglaophyton as unplaced in Horneophytopsida (alternative name: Langiophytopsida), while Novikoff & Barabasz-Krasny, 2015 have Aglaophyton/Lyonophyton in fam. Lyonophytaceae in order Lyonophytales which, together with Langiophytales, make up the class Langiophytopsida, sole member of the Division Langiophytophyta. I can always update IRMNG to follow this treatment if it seems preferable.

Where the "eophytes" would sit relative to this, I do not know, however others might...

Regards - Tony

Tony Rees, New South Wales, Australia
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On Sat, 1 Apr 2023 at 09:16, Kenneth Kinman <kinman at hotmail.com<mailto:kinman at hotmail.com>> wrote:
Dear All,
      I wasn't going to comment on this thread, but I never liked the name Eophytidae and I like the name "Eophyta" even less (especially at Phylum level).  If you think about it, the only true "eophytes" are cyanophyta/cyanobacteria.
       Anyway, since IRMNG already includes genus Aglaophyton in Class Horneophytopsida, which probably makes it paraphyletic, why not place Subclass Eophytidae in that class as well?  Then you don't have to worry about changing the name at all.
                              ------------Ken Kinman
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From: Taxacom <taxacom-bounces at lists.ku.edu<mailto:taxacom-bounces at lists.ku.edu>> on behalf of Tony Rees via Taxacom <taxacom at lists.ku.edu<mailto:taxacom at lists.ku.edu>>
Sent: Wednesday, March 29, 2023 5:00 PM
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Subject: Taxacom: Question to botanical experts RE ICNafp Code - rank changes for descriptive names, with changed termination

Dear all,

I am seeking advice as to how a descriptive name above the rank of family
can be re-used, with a change in termination (to signify changed rank) with
or without a change in cited authorship and/or "stat. nov." - the latter
presumably requiring a formal publication.

The name in question is Eophytidae Edwards, Morris, Axe, Duckett, Pressel &
Kenrick, 2021, erected as a subclass within class Embryopsida, consistent
with the treatment of land plants by Chase & Reveal, 2009, except that
"Embryopsida" replaces the latter's "Equisetopsida".

This name is erected at a similar rank to bryophytes, marchantiophytes atc.
which are at equivalent rank i.e. subclass in the Chase & Reveal treatment.
However in other treatments such as that of Ruggiero et al., 2015, followed
in my own IRMNG system and elsewhere, bryyophytes, etc. are treated as
separate phyla within Embryophyta, which is represented as a superphylum.

So my requirement is to treat the "eophytes" (presently Eophytidae) as a
separate phylum, e.g. "Eophyta", where it will sit alongside
Anthocerotophyta, Bryophyta, Charophyta and so on. Note here, "Eophytidae"
is a "descriptive name" per the ICNafp (not based on a genus) so falls
under the provisions of Arts. 16 and 6.10, thus:
-------------
Article 16.1 Names above the rank of family
"… descriptive names … may be used unchanged at different ranks"
-------------
Article 6.10 Note 3
Note 3. A descriptive name (Art. 16.1(b)) used at a rank different from
that at which it was first validly published is not a name at new rank
because descriptive names may be used unchanged at different ranks.
-------------
So one reading of this is that Eophytidae may be recast as "Eophyta" (with
the same circumscription) without a change of authorship, and not requiring
a published "stat. nov.". However the alternative reading would be that a
published change of authorship would be needed, plus a "stat. nov.", since
although the initial portion of the name is unchanged, the termination is -
in other words the name is not "unchanged" in the sense of Art. 16.1.

Thoughts, advice appreciated.

Tony Rees, New South Wales, Australia
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